Riodinidae

Mariposas de Bolivia

sitio 55 

 first update d.  4  february 2005

  last update d.  25 december 2021

Redactor: Lars Andersen

bajo construccion

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Metalmarks

Riodinidae

Subfamily Euselasiinae

Tribe Euselasiini

Athena Euselasia  (Euselasia athena). Coroico, Yungas, Bolivia. d. 21 januar 2006. Fotograf: Lars AndersenAthena Euselasia  (Euselasia athena). Coroico 1800 m.a., Yungas. d. 23 january 2008. Photographer: Lars AndersenAthena Euselasia  (Euselasia athena). Coroico 1800 m.a., Yungas. d. 23 january 2008. Photographer: Lars AndersenAthena Euselasia  (Euselasia athena). Coroico 1800 m.a., Yungas. d. 24 january 2008. Photographer: Lars Andersen

Euselasia eustola eustola (Stichel, 1919)

Eumenes Euselasia (Euselasia eumenes). Coroico, Yungas, d. 24 February 2009. Photographer; Lars AndersenEumenes Euselasia (Euselasia eumenes). Coroico, Yungas, d. 24 February 2009. Photographer; Lars AndersenEumenes Euselasia (Euselasia eumenes). Coroico, Yungas, d. 24 February 2009. Photographer; Lars Andersen

Euselasia eusepus (Hewitson, 1853)

Subfamily Riodininae

Tribe Mesosemiini

Judicialis Eyemark (Mesosemia judicialis). Caranavi, Yungas d. 17 January 2009. Photographer; Peter MøllmannMesosemia nerine (Stichel, 1909). Teoponte Mayaya, Caranavi, Yungas, Bolivia January 2015. Fotographer; Peter Møllmann

Mesosemia thymetus umbrosa (Stichel, 1909) & Mesosemia nerine (Stichel, 1909)

Coroico 1900 m.h. d. 11 February 2012. Photographer Lars AndersenCroesus Eyemark (Semomesia croesus). Caranavi, Yungas d. 17 January 2009. Photographer; Peter MøllmannMinos Eyemark   (Mesosemia minos). Caranavi, Yungas. d. 15 february 2007. Photographer: Lars Andersen

Mesosemia sp, Semomesia croesus siccata (Stichel, 1909) & Mesosemia minos (Hewitson, 1859)

Coroico, Yungas, d. 22 February 2009. Photographer; Lars AndersenCoroico, Yungas, d. 22 February 2009. Photographer; Lars AndersenCoroico, Yungas, d. 22 February 2009. Photographer; Lars AndersenCoroico, Yungas, d. 22 February 2009. Photographer; Lars Andersen

Mesosemia metuana glaucoma (Stichel, 1909)?

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Immature stages of the Rubiaceae-feeding metalmark butterflies (Lepidoptera: Riodinidae), and a new function for the tentacle nectary organs

November 2018

Gregory J. Nielsen and Lucas A. Kaminski; Metalmark butterfly (Riodinidae) diversity is heavily concentrated in the Neotropical lowland forests but despite their prevalence basic information on immature stages is still scarce.
Here we describe the life cycle of seven taxa in the
Rubiaceae-feeding section of the Mesosemiina, including three Mesosemia Hübner, [1819], three Leucochimona Stichel, 1909, and the first information for Semomesia Westwood, 1851. Immature stage morphology of Mesosemia cippus Hewitson, 1859 is described in detail through scanning electron microscopy.
Generally, eggs are laid singly and caterpillars are folivorous with a cryptic green coloration except for M. cippus which has gregarious caterpillars with a bright yellow color pattern.

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Caranavi, Yungas. d. 22 february 2007. Photographer: Lars Andersen

Actoris metalmarks, Napaea actoris (Cramer, 1776)

Tribe Eurybiini

Bluewing Sheenmark (Eurybia lycisca), Hotel Esmeralda, Coroico, Yungas, Bolivia d. 19 Januar 2005. Fotograf: Lars Andersen

Underleaf, Eurybia patrona promota (Stichel, 1910)

Tribe Riodinini

Caranavi, Yungas. d. 30 january 2007. Photographer: Lars AndersenCaranavi, Yungas. d. 30 january 2007. Photographer: Lars AndersenCaranavi, Yungas, Bolivia. d. 4 februar 2006. Fotograf: Lars AndersenSuapi, Yungas, Bolivia. D. 20 February 2009. Photographer: Lars AndersenRio Zongo,  between Caranavi and Guarnay, Yungas. d. 31 January 2009. Photographer: Lars AndersenRio Zongo,  between Caranavi and Guarnay, Yungas. d. 2 February 2008. Photographer: Lars AndersenLyropteryx apollonia, Challa, Yungas, i Januar 2005. Fotograf: Lars AndersenCaranavi, Yungas, Bolivia. d. 4 februar 2006. Fotograf: Lars AndersenSuapi, Yungas, Bolivia. D. 20 February 2009. Photographer: Lars Andersen

Blue-rayed Metalmark, Lyropteryx apollonia (Westwood, 1851)

Between Cascadas de San Jacinto Rio and Coroico. Yungas. d. 26 january 2007. Photographer: Lars AndersenBetween Cascadas de San Jacinto Rio and Coroico, 2000 m.a.. Yungas. d. 26 january 2007. Photographer: Lars AndersenBetween Cascadas de San Jacinto Rio and Coroico, 2000 m.a.. Yungas. d. 26 january 2007. Photographer: Lars AndersenBetween Cascadas de San Jacinto Rio and Coroico, 2000 m.a.. Yungas. d. 26 january 2007. Photographer: Lars Andersen

Black-rayed Metalmark, Lyropteryx terpsichore (Westwood, 1851)

Yolosa. date 25 January 2007. Photographer: Lars AndersenCaranavi, Yungas, Bolivia. d. 2 februar 2006. Fotograf: Lars AndersenSuapi, Yungas, Bolivia. D. 21 February 2009. Photographer: Lars AndersenCaranavi, Yungas, Bolivia. d. 4 februar 2006. Fotograf: Lars AndersenCaranavi, Yungas, Bolivia. d. 1 februar 2006. Fotograf: Lars AndersenThe old railroad / Kori Wayku inca trail, Yungas, elev. 2000 m. 27 January 2009. Photographer: Lars Andersen Cloudy mountainforest; The old railroad between Coroico and Coripata,Yungas. d. 24  February 2008. Photographer: Prem RoyCloudy mountainforest; The old railroad between Coroico and Coripata,Yungas. d. 24  February 2008. Photographer: Prem RoyThe old railroad / Kori Wayku inca trail, Yungas, elev. 2000 m. 27 January 2009. Photographer: Lars Andersen

Bellona Metalmark, Necyria bellona bellona (Westwood, 1851)

Challa,  between Caranavi and Coroico, Yungas. d. 25 February 2008. Photographer: Lars Andersen

Glasswing Metalmark, Ithomeis aurantiaca lauronia  (Schaus, 1902)

Celina Metalmark (Chimastrum celina). Caranavi, Yungas, Bolivia. d. 3 februar 2006. Fotograf: Lars AndersenCelina Metalmark (Chimastrum celina). Suapi, Yungas, Bolivia. D. 20 February 2009. Photographer: Lars AndersenCelina Metalmark (Chimastrum celina). Suapi, Yungas, Bolivia. D. 20 February 2009. Photographer: Lars Andersen

White-rayed Metalmark, Brachyglenis esthema esthema (C. Felder & R. Felder, 1862)

Erota Metalmark (Notheme erota). Rio Zongo,  between Caranavi and Guarnay, Yungas. d. 17 February 2009. Photographer: Lars AndersenErota Metalmark (Notheme erota). Rio Broncini, Caranavi, Yungas, Bolivia d. 13 Januar 2005. Fotograf: Lars Andersen

Tawny Metalmark, Notheme erota diadema (Stichel, 1910)

Albertus Metalmark (Monethe albertus). Caranavi, Yungas. d. 29 january 2007. Photographer: Lars Andersen

Albertus Metalmark, Monethe albertus (C. Felder & R. Felder, 1862)

Theodora Metalmark (Chalodeta theodora). Caranavi, Yungas. d. 30 january 2007. Photographer: Lars AndersenTheodora Metalmark (Chalodeta theodora). Caranavi, Yungas. d. 30 january 2007. Photographer: Lars AndersenTheodora Metalmark (Chalodeta theodora). Vagantes,  Yungas, Bolivia d. 21 januar 2005. Fotograf: Lars AndersenTheodora Metalmark (Chalodeta theodora). Yungas, Bolivia d. 15 Januar 2005. Fotograf: Lars Andersen

Theodora Metalmark, Chalodeta theodora (C. Felder & R. Felder, 1862)

Rio Coroico,  between Caranavi and Guarnay, Yungas. d. 18 February 2008. Photographer: Lars AndersenRio Coroico,  between Caranavi and Guarnay, Yungas. d. 18 February 2008. Photographer: Lars AndersenCaranavi, Yungas. d. 12 February 2009. Photographer: Lars AndersenCaranavi, Yungas. d. 12 February 2009. Photographer: Lars AndersenCaranavi, Yungas. d. 12 February 2009. Photographer: Lars AndersenThies is a undiscovery species! Caranavi, Yungas, Bolivia. february 2006. Photographer: Ib KreutzerCaranavi, Yungas. d. 12 February 2009. Photographer: Lars AndersenCaranavi, Yungas. d. 12 February 2009. Photographer: Lars AndersenCaranavi, Yungas. d. 12 February 2009. Photographer: Lars AndersenCaranavi, Yungas. d. 12 February 2009. Photographer: Lars Andersen

Orange-stitched Metalmark, Chalodeta chaonitis (Hewitson, 1866)

Caranavi, Yungas, Bolivia d. 13 Januar 2005. Fotograf: Lars Andersen

Chamaelimnas unknown species?

Caranavi, Yungas, Bolivia  d. 19 january 2010. Photographer; Lars AndersenCaranavi, Yungas. d. 30 january 2007. Photographer: Lars AndersenCaranavi, Yungas. d. 30 january 2007. Photographer: Lars AndersenCaranavi, Yungas. d. 30 january 2007. Photographer: Lars AndersenTocana, Yungas, Bolivia. d. 24 januar 2006. Fotograf: Lars AndersenCaranavi, Yungas. d. 29 January 2008. Photographer: Lars Andersen

Chamaelimnas splendens (Grose-Smith, 1902)

Caranavi, elev. 600 m. d.  26 January 2012. Photographer: Lars AndersenOrange-stitched Metalmark (Chalodeta chaonitis). Rio Zongo,  between Caranavi and Guarnay, Yungas. d. 31 January 2009. Photographer: Lars AndersenOrange-stitched Metalmark (Chalodeta chaonitis). Rio Zongo,  between Caranavi and Guarnay, Yungas. d. 31 January 2009. Photographer: Lars AndersenOrange-stitched Metalmark (Chalodeta chaonitis). Vagantes, Yungas, Bolivia d. 22 januar 2005. Fotograf: Annelie Nielsen

Gynaea Metalmark, Detritivora gynaea (Godart, 1824)

Puna, Coroico 1900 m.h. d. 13 february 2012. Photographer Lars AndersenPuna, Coroico 1900 m.h. d. 13 february 2012. Photographer Lars AndersenCoroico, Yungas, d. 22 February 2009. Photographer; Lars AndersenPuna, Coroico 1900 m.h. d. 13 january 2012. Photographer Lars Andersen

Calephelis laverna (Godman & Salvin, 1886)

Schaus' Metalmark (Calephelis schausi). Caranavi, Yungas, Bolivia d. 14 januar 2005. Fotograf: Lars AndersenSchaus' Metalmark (Calephelis schausi). Tocana, Yungas, Bolivia. d. 24 januar 2006. Fotograf: Lars AndersenSchaus' Metalmark (Calephelis schausi). Vagantes, Yungas, Bolivia. d. 25 januar 2006. Fotograf: Lars AndersenSchaus' Metalmark (Calephelis schausi). Vagantes, Yungas, Bolivia. d. 25 januar 2006. Fotograf: Lars AndersenSchaus' Metalmark (Calephelis schausi). Caranavi, Yungas. d. 15 february 2007. Photographer: Lars Andersen

Calephelis aymaran (McAlpine, 1971)

Amarynthina Metalmark (Parcella amarynthina). Vagantes, Yungas, Bolivia. d. 25 januar 2006. Fotograf: Lars AndersenAmarynthina Metalmark (Parcella amarynthina), Rio Broncini, Caranavi, Yungas, Bolivia d.13 januar 2005. Fotograf: Lars AndersenAmarynthina Metalmark (Parcella amarynthina). Vagantes, Yungas, Bolivia. d. 25 januar 2006. Fotograf: Lars Andersen

Orange-banded Metalmark, Parcella amarynthina (C. Felder & R. Felder, 1862)

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Emeralds Metalmarks, Caria

Caranavi, Yungas, Bolivia d. 19 January 2010. Photographer; Lars Andersen

Green Mantle, Caria unknown species? Caranavi, Yungas, Bolivia d. 19 January 2010. Photographer; Lars Andersen

See more about these mysterious beautiful jewels on the side with Emeralds Metalmarks

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Fasciata Metalmark (Crocozona fasciata). Coroico, Yungas, Bolivia d. 1 januar 2005. Fotograf: Lars AndersenFasciata Metalmark (Crocozona fasciata). Coroico 1800 m.a., Yungas. d. 24 january 2008. Photographer: Lars Andersen

Crocozona fasciata clusia (Hewitson, 1866)

Fasciata Metalmark (Crocozona fasciata). Hotel Esmeralda, Coroico, Yungas, Bolivia d. 17 januar 2005. Fotograf: Lars Andersen

Crocozona species?

Coecias Metalmark (Crocozona coecias). Caranavi, Yungas. d. 16 February 2009. Photographer: Lars AndersenCoecias Metalmark (Crocozona coecias). Caranavi, Yungas. d. 12 February 2009. Photographer: Lars AndersenCoecias Metalmark (Crocozona coecias). Caranavi, Yungas, Bolivia. d. 1 februar 2006. Fotograf: Lars AndersenCoecias Metalmark (Crocozona coecias). Bolinda, Caranavi, Yungas. d. 12 february 2007. Photographer: Lars AndersenCoecias Metalmark (Crocozona coecias). Bolinda, Caranavi, Yungas. d. 12 february 2007. Photographer: Lars Andersen

Crocozona coecias (Hewitson, 1866)

Bacaenis Metalmark  (Baeotis bacaenis). Caranavi, Yungas, Bolivia. d. 3 februar 2006. Fotograf: Lars AndersenBacaenis Metalmark  (Baeotis bacaenis). Caranavi, Yungas, Bolivia. d. 2 februar 2006. Fotograf: Lars AndersenBacaenis Metalmark  (Baeotis bacaenis). Caranavi, Yungas, Bolivia. d. 2 februar 2006. Fotograf: Lars AndersenBacaenis Metalmark  (Baeotis bacaenis). Caranavi, Yungas. d. 3 february 2007. Photographer: Lars AndersenBacaenis Metalmark  (Baeotis bacaenis). Yolosa, Yungas, Bolivia d. 10 Januar 2005. Fotograf: Lars AndersenBacaenis Metalmark, Baeotis bacaenis. Suapi, Yungas, Bolivia. D. 21 February 2009. Photographer: Lars Andersen

Baeotis elegantula (Hopffer, 1874)

Baeotis creusis. Kori Wayku inca trail elev. 2000 m. d.  14 December 2014. Photographer:  Peter Møllmann

Baeotis creusis (Hewitson, 1874)

Caranavi, Yungas, Bolivia d. 19 January 2010. Photographer; Lars AndersenCaranavi, Yungas. d. 3 february 2007. Photographer: Lars AndersenCaranavi, Yungas. d. 3 february 2007. Photographer: Lars AndersenD'Abrera Pigmy, Baeotis staudingeri (D'Abrera, 1994). Garrapatuni Caranavi, Yungas, Bolivia 2th january 2015. Fotograf; Peter Møllmann

D'Abrera Pigmy, Baeotis staudingeri (D'Abrera, 1994)

Lasaia agesilas. Caranavi, Yungas. d. 19 february 2007. Photographer: Lars AndersenCaranavi, Yungas. d. 6 February 2009. Photographer: Lars AndersenCaranavi, Yungas. d. 6 February 2009. Photographer: Lars AndersenVagantes, Yungas, Bolivia. d. 25 januar 2006. Fotograf: Lars Andersenblue lasaia, Lasaia moeros,Tocana, Yungas, i Januar 2005. Fotograf: Lars AndersenTocana, Yungas, Bolivia. d. 20 januar 2006. Fotograf: Lars AndersenCaranavi, Yungas. d. 7 February 2009. Photographer: Lars AndersenCaranavi, Yungas. d. 6 February 2009. Photographer: Lars Andersen

Glittering Sapphire or Shining-blue Lasaia, Lasaia agesilas (Latreille, 1809)

Lasaia arsis. Caranavi, Yungas. d. 6 february 2007. Photographer: Lars AndersenCaranavi, Yungas, Bolivia. D. 25 January 2010. Photographer: Lars Andersen Caranavi, elev 870 m. Yungas. d. 8 january 2016. Photographer; Peter  Møllmann

Cat's-eye Sapphire, Lasaia arsis (Staudinger, 1887)

The genus Sapphire Metalmarks, Lasaia (H. Bates, 1868) contains arround 14 species, all of which are found exclusively in the neotropics. They are small butterflies, averaging about 28 to 32 mm in wingspan. Males have extremely reflective wing scales, shimmering in metallic turquoise, blue or steely grey according to species. Females are generally a dull earthy brown colour. Both sexes have a similar pattern of black spots.

Males are often seen along the sandbanks with others butterflies along small streams, where they suck minerals and ammonia, as well as the droppings from mammals.

Females are rarely seen, and lives mostly up in the bushes and trees of Mimose, Albizia species. Pea, Fabaceae family which is the foodplant for the caterpillar.

Lars Andersen and Peter Møllmann has so far seen three species in Bolivia: Glittering Sapphire, Lasaia agesilas (Latreille, 1809). Cat's-eye Sapphire, Lasaia arsis (Staudinger, 1887) & Kenneth's Sapphire, Lasaia kennethi (Weeks, 1901). But there are certainly several species?

Sapphire Metalmarks species you maybe can expect to find in Bolivia are: Variegated Sapphire, Lasaia meris (Stoll, 1781). Bolivian Sapphire, L. moeros (Staudinger, 1888). L. aerugo (Clench, 1972). Clench's Sapphire, L. pseudomeris (Clench, 1972). Oily Sapphire, L. ouleus (Godman, 1903) and L. incoides (Schaus, 1902).

Lasaia kennethi. Caranavi, Yungas. d. 2 february 2007. Photographer: Lars AndersenCaranavi, Yungas, Bolivia. d. 30 januar 2006. Fotograf: Lars AndersenTocana, Yungas, Bolivia. d. 24 januar 2006. Fotograf: Lars AndersenTocana, Yungas, Bolivia. d. 24 januar 2006. Fotograf: Lars AndersenKenneth's Metalmark, Lasaia kennethi. Suapi, Yungas, Bolivia. D. 21 February 2009. Photographer: Lars Andersen

Kenneth's Sapphire or Kenneth's Lasaia, Lasaia kennethi (Weeks, 1901)

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Caranavi, Yungas. d. 3 February 2009. Photographer: Lars AndersenCaranavi, Yungas. d. 8 February 2009. Photographer: Lars AndersenRio Zongo,  between Caranavi and Guarnay, Yungas. d. 2 February 2008. Photographer: Lars AndersenAmrynthis menena, Rio Broncini, Caranavi, Yungas, Bolivia d. 13 Januar 2005. Fotograf: Lars AndersenCaranavi, Yungas, Bolivia. d. 1 februar 2006. Fotograf: Lars AndersenCaranavi, Yungas, Bolivia. d. 1 februar 2006. Fotograf: Lars Andersen

Red-barred Amarynthis, Amarynthis meneria (Cramer, 1776)

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Black Metalmark, Melanis

Electron Pixie, Melanis electron (Fabricius, 1793). Caranavi, Yungas. d. 12 february 2007. Photographer: Lars Andersen

Electron Pixie, Melanis electron (Fabricius, 1793). Caranavi, Yungas, Bolivia d. 12 february 2007. Photographer; Lars Andersen

See more about these black jewels on the side with Black Metalmark

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Tribe Symmachiini

Xynias lithosina cynosema (Hewitson, 1874).Quijarro-Caranavi, Yungas. d. 16 february 2007. Photographer: Lars Andersen

Xynias lithosina cynosema (Hewitson, 1874)

Clonia Metalmark, Esthemopsis clonia (C. Felder & R. Felder, 1865). Coroico, Yungas, d. 1 Januar 2005. Fotograf: Lars Andersen

Clonia Metalmark, Esthemopsis clonia (C. Felder & R. Felder, 1865)

En metalvinge fra Coroico, Yungas, d. 31 december 2004. Fotograf: Lars Andersen

Accused Metalmark, Symmachia accusatrix (Westwood, 1851)

Tribe Helicopini

Anteros kupris aureocultus. Kori Wayku inca trail, Yungas, Bolivia. d. 2 Febrary 2010. photographer; Lars Andersen

Great Jewelmark, Anteros kupris aureocultus (Stichel, 1909)

Forest near the Viluny antenna and Caranavi Valley 800 to 1200 m., Yungas, Bolivia december 13 - 20, 2021. Photographer;  Gottfried SiebelPrincipalis Jewelmark, Anteros principalis (Hopffer, 1874). Caranavi Highlands, Yungas, Bolivia  january 28, 2019. Photographer; Peter MøllmannPrincipalis Jewelmark, Anteros principalis (Hopffer, 1874).Vagantes, Coroico, Yungas, Bolivia november 4, 2017. Photographer; Hiroaki Takenouchi

Principalis Jewelmark,
Anteros principalis (Hopffer 1874).
Caranavi 1200 m., Yungas, Bolivia
december 13 - 20, 2021.
Photographer; 
Gottfried Siebel

Principalis Jewelmark,
Anteros principalis (Hopffer, 1874).
Caranavi Highlands, Yungas, Bolivia  
january 28, 2019.
Photographer; Peter Møllmann

Principalis Jewelmark,
Anteros principalis (Hopffer, 1874).
Coroico, Yungas, Bolivia
november 4, 2017.
Photographer; Hiroaki Takenouchi

Principalis Jewelmark, Anteros principalis (Hopffer, 1874).

The adults are elusive, and usually encountered singly, in the vicinity of rivers or streams. They normally rest on the upper surface of leaves, and always hold their wings erect when settled.

This species is found in undisturbed rainforest habitats at altitudes between 0-1100m.

The caterpillar folds the edge of a leaf into a tube and lives within. The foodplants include
Vochysia (Vochysiaceae) and Miconia (Melastomataceae).

Is distributed on the eastern slopes of the Andes from Colombia, Ecuador to Peru and Bolivia.

There are 22 species in the genus Anteros, distributed variously from Mexico to Bolivia. The genus is closely allied to Sarota and Helicopis in Riodinidae family, and like them is characterised by having silvery markings on the underside wings, and very "furry" legs.

Butterflies of America 
and the Interactive Listing of American Butterflies

Caranavi, Yungas, Bolivia. d. 1 februar 2006. Fotograf: Lars AndersenCaranavi, elev. 1400 m. d.  27 January 2012. Photographer: Lars Andersen

Starry Night Metalmark, Echydna punctata (C. Felder & R. Felder, 1861)

En metalvinge fra Challa, Yungas, i Januar 2005. Fotograf: Lars AndersenRio Broncini, Caranavi, Yungas, Bolivia d. 13 Januar 2005. Fotograf: Lars AndersenCaranavi, Yungas, Bolivia. d. 30 januar 2006. Fotograf: Lars Andersen

Stichel's Blue Emesis, Emesis orichalceus (Stichel, 1916)

Agularis Metalmark (Emesis angularis). Kori Wayku inca trail, Yungas, Bolivia d. 1 february 2010. Photographer; Lars AndersenCamino de Muerte elev. 2600 m. Yungas, Bolivia d.  18 January 2012. Photographer: Lars AndersenKori Wayku inca trail elev. 2000 m. d.  15 January 2012. Photographer: Lars Andersen

Emesis angularis (Hewitson, 1870)

Caranavi, Yungas. d. 12 February 2009. Photographer: Lars AndersenTocana, Yungas, Bolivia. d. 20 januar 2006. Fotograf: Lars AndersenCaranavi elev. 910 m. d.  23 January 2012. Photographer: Lars Andersen

Dark Emesis, Emesis ocypore (Geyer, 1837)

Caranavi 800 m. d.  14 December 2014. Photographer:  Peter Møllmann

Silky Emesis, Emesis brimo progne (Godman, 1903)

Rio Zongo,  between Caranavi and Guarnay, Yungas. d. 26 January 2008. Photographer: Lars AndersenCaranavi, Yungas. d. 8 February 2009. Photographer: Lars AndersenCaranavi, Yungas. d. 3 february 2007. Photographer: Lars AndersenCaranavi, Yungas, Bolivia. d. 30 januar 2006. Fotograf: Lars AndersenCaranavi, Yungas, Bolivia. d. 30 januar 2006. Fotograf: Lars AndersenMandana Metalmark (Emesis mandana), Rio Broncini, Caranavi, Yungas, Bolivia d. 13 Januar 2005. Fotograf: Lars Andersen

Great Emesis, Emesis mandana (Cramer, 1780)

Caranavi, Yungas, Bolivia d.11 januar 2005. Fotograf: Lars AndersenTocana, Yungas, Bolivia. d. 24 januar 2006. Fotograf: Lars AndersenCaranavi d. 29 january 2008. Photographer; Lars Andersen

Orange-barred Emesis, Emesis cypria (Felder & felder, 1861) and

Caranavi, Yungas. d. 7 february 2007. Photographer: Lars AndersenCaranavi, Yungas. d. 7 february 2007. Photographer: Lars Andersen

Emesis temesa (Hewitson, 1870)

Caranavi, Yungas. d. 12 February 2009. Photographer: Lars AndersenCaranavi, Yungas. d. 12 february 2007. Photographer: Lars AndersenCaranavi, Yungas. d. 1 February 2008. Photographer: Lars Andersen

White-marked Emesis, Emesis fastidiosa (Ménétriés, 1855)

Astraeodes areuta. Caranavi, Yungas, Bolivia. d. 2 februar 2006. Fotograf: Lars AndersenAstraeodes areuta. Rio Zongo,  between Caranavi and Guarnay, Yungas. d. 26 January 2008. Photographer: Lars Andersen

Astraeodes areuta (Westwood, 1851)

Tribe Nymphidiini

Zygia Metalmark (Lemonias zygia). Caranavi, Yungas. d. 7 february 2007. Photographer: Lars AndersenZygia Metalmark (Lemonias zygia). Caranavi, Yungas. d. 7 february 2007. Photographer: Lars Andersen

Lemonias zygia (Hübner, 1807)

Caranavi, Yungas. d. 13 february 2007. Photographer: Lars Andersen

Molpe Metalmark, Juditha caucana (Stichel, 1911)

Menander menander. Caranavi 605 m. Yungas, Bolivia d.  18 January 2014. Photographer:  Ole Andersen

Menander Metalmark, Menander menander thallus (Stichel, 1911)

Caranavi, Yungas. d. 6 February 2009. Photographer: Lars Andersen

Zelotaea leucotopus (Stichel, 1911)

Emylius Metalmark (Calospila emylius crispinella). Caranavi, Yungas d. 8 January 2012. Photographer; Peter MøllmannCaranavi, Yungas. d. 31 January 2008. Photographer: Lars Andersen

Calospila emylius crispinella (Stichel, 1911)

Caranavi, Yungas, Bolivia. d. 2 februar 2006. Fotograf: Lars Andersen

Nymphidium ascolia (Hewitson, 1853)

Caranavi, Yungas. d. 3 February 2009. Photographer: Lars AndersenCaranavi, Yungas. d. 4 february 2007. Photographer: Lars Andersen

Nymphidium carmentis (Stichel, 1910)

Tribe Stalachtini


Black Lace, Chorinea faunus. Rio Rufus, Taipiplaya, Caranavi, Yungas. d. 8 February 2009. Photographer: Lars Andersen

Swordtails - Tribe Riodinini

sitio 57

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METALMARKS part1 from Peter Møllmann on Vimeo.


METALMARKS part3 from Peter Møllmann on Vimeo.

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Introduction to Metalmarks

Riodinidae are a pantropical family, with the majority of species occurring in the Amazonas and Andeans eastside. There are around 1300 to 1325? known species of Riodinidaein the neotropical region, from Mexico to Argentina.

Other Names for Riodinidae Grote 1895

  • Erycinidae Swainson 1827 (see nomenclature)
  • Vernacular Names: Metalmarks

From TREE of LIFE Web project!

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  Papilionidae
  Pieridae

Dayflying Butterflies family group: Papilionoidea :

Lycaenidae   Riodinidae:
  Nymphalidae

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Riodinidae:

  Euselasslini
Euselasiinae: Stygini
  Corrachiini
  Mesosemiini
  Eurybiini
  Riodinini
Riodininae: Symmachiini
  Helicopini
  Nymhidiini
  Stalachtini
  Riodininae- incertae sedis
  Nemobiinae

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dont use this pictures without permission from: 

photographer © Lars Andersen


Mariposas de Bolivia 

Enero 2009

por Peter Møllmann y Lars Andersen


link to Neotropical Butterflies of Will Carter and Kim Garwood, Metalmarks


Esmeralda hotel, Coroico, Yungas